The sooty falcon (Falco concolor) is a medium-sized falcon breeding from northeastern Africa to the southern Persian Gulf
region. The word sooty means to be covered in soot (ash), and is used
to describe the color of the Sooty Falcon. Hence, the falcon gets its
name from its color, the color of soot.
The Sooty Falcon is part of the order Falconiformes, family Falconidae and genus Falco. [2] The sooty falcon belongs to the hobby group, a rather close-knit number of similar falcons often considered a subgenusHypotriorchis. Eleonora's falcon is sometimes considered its closest relative, but while they certainly belong to the same clade, they do not seem to be sister species.[3][4]
This is an elegant bird of prey, 32–37 cm long with a 78–90 cm wingspan. It is shaped like a large hobby or a small Eleonora's falcon,
with its long pointed wings, long tail and slim body. The adults are
blue-grey, and lack the black underwing coverts of the Eleonora's
falcon. The young bird is like a large juvenile hobby, or small juvenile
Eleanora's falcon. Its dark trailing edge to the wings and tail
distinguish it from the former species, and it lacks the underwing
contrast caused by the dark coverts of the larger falcon.[5]
This species is distributed across Southwest Asia and North Africa. [6] This species breeds on islands and coastal or desert cliffs from Libya to Pakistan). It is a long-distance migrant, wintering in east Africa and south to Madagascar and south-eastern South Africa.
Increasingly regular sightings and a lack of historical records
suggests that the wintering range has expanded south in recent decades.[5] It is a rare vagrant north of its breeding range.
The sooty falcon eats mainly birds (e.g. Sandgrouse), but it will take large insects, such as dragonflies,
which are transferred from talons to beak and eaten in flight. They
have also been known to eat lizards and crabs. During their breeding
season they exclusively feed on small birds. [6]
During
winter the falcons hunt alone, in groups or even as flocks of up to 15
individuals. These hunting flocks tend to perch on trees and feed on
swarming insects. During the breeding, summer, season males do the
majority of the hunting. Four hunting techniques are employed: (1)
Diving from a perch. The falcons spot prey from their perch and then
dive bomb and catch it. (2) Flushing. Flushing is a technique used in
which falcon individuals fly low over bushes or trees in an attempts to
flush other birds out, after which they catch them. (3) Flying up to
come down. When a falcon spots a flying prey bird they try to fly
further above it, and then dive down to intercept the prey. (4) Survey
flight. Individuals circle areas at great heights in hopes of spotting a
prey item. Once prey is spotted the falcon flies down below the prey,
and then turns back up into the prey's flight path to catch it. [6]
Sooty
Falcons lay eggs during mid-summer, and unlike most other falcons,
occasionally nest in colonies. The falcons breed in the arid deserts of
Southwest Asia and North Africa during the summer months, which is
remarkable due to high summer temperatures. They breed in three types of
arid environment: (1) small islands in the Red Sea, (2) mountainous
deserts and (3) in deserts with rocky outcrops. It nests on a ledge or
on rocks, laying up to four eggs.[5]
While the falcons tend to nest on high cliffs, they have also been
found nesting between rocks or on the ground. Nests are scraped into
soft limestone, chalk or sandstone cliffs, and face away from the sun to
provide shelter from its rays. Nest also function as perches from which
defense against predators is mounted. [6]
Birds begin to display and mate during April and May, with eggs
being laid by July-August. Egg incubation lasts approximately 27-29
days. Incubation is predominantly performed by the female, while the
male only taking shifts for 1-7 minutes while the female sunbathes.
Males hunt and bring food to the female and chicks once they have
hatched. During the incubation period males provide females with 3-4
birds per day. Excess food is cached on shaded ledges outside the nest.
Doing this ensures that food is regulated and that it is available when
it is scarce. It also keeps the nests clean, and prevents parasites and
pests from being attracted to the nests. The young falcons remain in the
nest for 32-38 days, after which they still remain near the nest for a
further 2-3 weeks. During this period the parents still take care of
them, but thereafter the level of parental care begins to decrease. [6]
Birds nesting on islands in the Sea of Oman have a mean clutch
size of 2.83 and brood size of 2.11, with 12% of nests failing at the
egg or nesting stage.[7]
The
falcons migrate from their breeding grounds in Southwest Asia and North
Africa to winter in Eastern and South-eastern Africa. The journey takes
approximately 13 days and covers a distance of 5500 kilometers. The
falcons start their migration during the night but tend to travel only
during the day. They stop-over at rest spots along Eastern Africa, spots
where there are potential water sources. [8]
It was formerly classified as Least Concern by the IUCN, but has recently been shown to be rarer than formerly believed. Consequently, it was uplisted to Vulnerable status in 2017.[1] It is estimated that their population ranges between 2800 and 4000 individuals and that these numbers are declining. [6]
The breeding population of Sooty falcons on the islands of northern
Oman is in decline, with human disturbance on accessible islands a
likely cause, as falcons breeding close to beaches have lower nesting
success than those breeding further away.[9] Hence, mitigating human disturbances and protecting natural habitats is of critical conservation concern to the Sooty Falcon.
The wedge-tailed eagle (Aquila audax) also known as the eaglehawk, is the largest bird of prey in the continent of Australia. It is also found in southern New Guinea to the north and is distributed as far south as the state of Tasmania.[1]
Adults of the species have long, broad wings, fully feathered legs, an
unmistakable wedge-shaped tail, an elongated upper mandible, a strong
beak and powerful feet.[4] The wedge-tailed eagle is one of 12 species of large, predominantly dark-coloured booted eagles in the genus Aquila
found worldwide. Genetic research has clearly indicated that the
wedge-tailed eagle is fairly closely related to other, generally large
members of the Aquila genus.[5] A large brown-to-black bird of prey, it has a maximum reported wingspan of 2.84 m (9 ft 4 in) and a length of up to 1.06 m (3 ft 6 in).[6]
The wedge-tailed eagle is one of the world's most powerful avian predators.[8] Although a true generalist, which hunts a wide range of prey, including birds, reptiles and, rarely, other taxa, the species is, by and large, a mammal predator.[8][9] The introduction of the European rabbit (Oryctolagus cuniculus) has been a boon to the wedge-tailed eagle and they hunt these and other invasive species in large volume, although the wedge-tailed eagle otherwise generally lives off of marsupials, including many surprisingly large macropods. Additionally, wedge-tailed eagles often eat carrion, especially while young.[7][8][10] The species tends to pair for several years, possibly mating for life.[8]
Wedge-tailed eagles usually construct a large stick nest in an
ample tree, normally the largest in a stand, and typically lay two eggs,
although sometimes one to four.[7] Usually, breeding efforts manage to produce one or two fledglings which, after a few months more, tend to disperse widely.[9] Nesting failures are usually attributable to human interference, such as logging
activity and other alterations, which both degrade habitats and cause
disturbances. The species is known to be highly sensitive to human
disturbance at the nest, which may lead to abandonment of the young.[11]
Although historically heavily persecuted by humans through poisoning and shooting, mostly for alleged predation on sheep,
wedge-tailed eagles have proved to be exceptionally resilient, and
their numbers have quickly rebounded to being similar or even higher
numbers than before European colonisation, thanks in part to humans inadvertently providing several food sources, such as rabbits and a large volume of roadkill.[1][7][12]
An adult wedge-tailed eagle at Lake Burrumbeet in flight, notably dark and blackish colour.
A young wedge-tailed eagle perched in Birdsville Track with an unusual amount of pale colour showing due likely to moult.
The species was first described in 1801 by the English ornithologist John Latham, under the binomial nameVultur audax.[13] At one time, the wedge-tailed eagle was classified in it is own monotypical genus Uroaetus, perhaps due to its unique form.[6]
Today, the genus Vultur is used only for a completely unrelated bird of the New World vulture family, the Andean condor (Vultur gryphus).[14] The specific scientific name for the species, audax, is derived from the Latinaudax,
meaning "bold", indicative of their perceived disposition, perhaps when
hunting, although the species is, in general, highly wary, and even
timid, around humans.[8]
However, the species is quite similar in many aspects of its
morphology, appearance, behaviour and life history, to other species in
the Aquila genus.[6][10][15] The eagles of the Aquila genus are part of the subfamily Aquilinae, within the larger Accipitridae
family. The subfamily is commonly referred to as booted eagles or
sometimes as true eagles. Those species may be distinguished from most
other accipitrids by the feathering covering their legs, regardless of
distribution. With some 39 or so species, the Aquilinae is present on
every continent except Antarctica.[14][15]
By a variety of phylogenetic testing, largely via Mitochondrial DNA and Nuclear DNA genes, it has been determined that the wedge-tailed eagle clusters with certain other Aquila eagles. The species found to share the most genetic similarities is the Verreaux's eagle (Aquila verreauxii) of Africa. However, the Gurney's eagle (Aquila gurneyi), a mostly allopatric but outwardly fairly similar eagle, is clearly a very close relation of the wedge-tailed eagle and the two are likely sister species, most probably originating from the same radiation across the Indo-Pacific region.[5][7][8][16]
The wedge-tailed, Gurney's and Verreaux's eagles form a clade or a species complex with the well-known golden eagle (Aquila chrysaetos),
the most widely distributed species in the entire accipitrid family, as
well as outwardly dissimilar (smaller and paler-bellied yet also
powerful) eagles like the Bonelli's eagle (Aquila fasciata), the African hawk-eagle (Aquila spilogaster) and the Cassin's hawk-eagle (Aquila africanus), the latter three having once been considered members of a different genus.[citation needed]
Beyond the aforementioned species, based on genetic testing, the four other Aquila
species, although outwardly similar to golden and wedge-tailed eagles,
being large, dark and brownish, with long wings, are thought to form a
separate clade, and are paraphyletic from the members of what can be called the golden eagle clade.[5][7][16][17] Other related outliers from outside the Aquila genus, are the small-to-mid-sized Clanga or spotted eagle species, and the widely found and quite small Hieraeetus eagles. One member of the latter genus contains the only other widely found Aquilinae eagle in Australia, the little eagle (Hieraaetus morphnoides).[5][16][17]
A. a. fleayi, mobbed by forest ravens in Tasmania
Two subspecies of wedge-tailed eagle are recognised.[18] However, the separation of the two subspecies has been called into question, largely because the reported differences in both size and coloration can be attributed to clinal variation, and some of the insular populations may still be at an intermediate stage of subspecific formation.[7][8][19]A. a. audax (Latham, 1801) – This subspecies resides in
the entire continent of Australia as well as in southern New Guinea. It
is the typical wedge-tailed eagle as subsequently described.[citation needed] A. a. fleayi (Condon & Amadon, 1954) – This race is endemic to Tasmania. The subspecies is named in honour of David Fleay, an Australian naturalist who was the first to propose the difference of the insular race.[8]A. a. fleayi differs from mainland wedge-tailed eagles mainly via size and colouring.[4] It is larger than the mainland eagle and is said to have particularly outsized talon dimensions compared to mainland eagles.[20]
Furthermore, it has a deep chocolate brown overall colour rather than
blackish, with a whitish buff colouring to the nape rather than
tawny-rufous feathers there.[4][8] The juvenile is altogether paler and sandier than an equivalent-aged wedge-tailed eagle on mainland Australia.[8][9]
Although the validity of the subspecies has been questioned, genetic studies have determined that there is no gene flow or introgression
between Tasmanian and other wedge-tailed eagles. Furthermore, the
insular race was likely formed by marine dispersals, a process
wedge-tailed eagles may continue to engage in despite usually avoiding
large bodies of water, albeit usually in narrower straits.[19][21]
In flight, the wedged tail is clearly visible. A wedge-tailed eagle foraging on the ground.
Wedge-tailed eagles are very large and quite lanky birds. They are characteristically black but can appear tar to charcoal brown, depending on lighting and individual variation.[9] They have a massive bill but possess a relatively small and rather flat head, with a long, almost vulturine neck. Furthermore, they are distinctive for their prominent carpals and baggy feathered trousers.[4]
The species tends to perch conspicuously on dead trees, telegraph
poles, rocks or, at times, the open ground. Between the bill size,
elongated shape and prominent shoulders, the species is highly
distinctive. While perched, their long wings extend down to a long and
markedly wedge-tipped tail.[4][9]
They have a large proportion of bare facial skin, which is
thought to be an adaptation to the warm climate rather than carrion
eating, because the non-carrion-eating Verreaux's eagle has similar facial feathering and the golden eagle eats carrion too.[8]
Against the blackish plumage, the tawny-rufous hackles on the neck,
forming a lanceolated shape, as well as the pale brown to rufous crissum, and narrow mottled grey-brown band across the greater wing coverts, all stand out well.[4][10] The sexes are indistinguishable by plumage.[4]
The juvenile is mainly darkish brown, with extensive rufous
feather edging, and a paler, fairly streaky head. Furthermore, the
juvenile has a lighter-brown crissum, and a light reddish-brown to
golden nape, with similar colouring extending sometimes to the back and
wing band. The wing band is considerably more prominent than those of
adults, extending to the median and sometimes the lesser coverts.
Rarely, a juvenile may be all dull black, lacking rufous edges or a wing
band.[4][7] Young eagles are much the same by the second through to the fourth years though they may be almost invariably visibly in moult
and with a narrowing wing band. They become darker around the fifth
year, with a red-brown nape and a still narrowing wing band. Full mature
plumage is not attained until the seventh or eighth year, although
sexual maturity can be considered as early as five.[4][7]
Adults have dark brown eyes, while juveniles usually have similar
but slightly darker eyes. Wedge-tailed eagles are typically creamy
white on the cere and feet, although those can be dull yellow, more so
in juveniles than adults.[4]
The wedge-tailed eagle has a unique moult process in that they moult
almost continuously and very slowly, and it might take three or more
years for an eagle of the species to complete a moult. Moults are
arrested only at times of famine, and happen gradually, so that they do
not impede the bird's flight or hunting capacities.[8][22]
In flight, wedge-tailed eagles appear as a very large, dark
raptor, with a protruding head, long and relatively narrow-looking
wings, more or less parallel edged when soaring and, most distinctly, a
long diamond-shaped tail. The shape is dissimilar to any other raptor in
the world.[4][23] Juveniles tend to be broader winged by comparison. The wingspan is around 2.2 times greater than the bird's total length.[4]
They tend to fly with rather loose but deep and powerful beats.
Wedge-tailed eagles spend much time sailing along, looking quite stable
and controlled even in strong winds. The species glides and soars on
upswept wings with long splayed primaries. The ample tail may be
upcurved, or "dished", at the edges.[4][24] The eagle often spreads its deep wing emarginations to reduce drag in high winds.[8][22]
Contrary to their superlative and controlled appearance once on the
wing, flight for wedge-tailed eagles can be a struggle even in normal
circumstances, unless from it is from a pinnacle or it is somewhat windy
and, within the forest, they may clamber about, with a "lack of grace",
to reach the canopy.[8] Gorged birds on the ground can be vulnerable, being practically grounded, which was an advantage historically to Aboriginal hunters.[8] Human gliders have encountered wedge-tailed eagles at more than 3,000 m (9,800 ft).[8]
The adult is all blackish on the wing but for the tawny-rufous
nape and greyish wing band (running less than a quarter of the way down
the wing's width). Little relieves the dark coloration below but the
pale brown to rufous crissum and the pale greyish bases to their flight feathers.[4][7]
Juvenile wedge-tailed eagles appear much browner although in general
are not dissimilar in pattern below though the body and wings relative
to adult. However, juveniles may show some paler mottling, of an
off-rufous colour. Meanwhile, the juvenile's tail and most flight
feathers are barred greyish which in turn contrast against the pale
based primaries with black tips. Above, the juvenile bears much paler
and more sandy rufous colour from the head to at least upper mantle and
along broad wing band (as well as more than half the wing width). The
lighter dorsal colour sometimes extends to much of the back and
scapulars. Rare individual juvenile eagles are dull black, without a
wing band or paler edges. With much variation in individuals, generally
as the young eagles age, the signature wing band shrinks incrementally
and, after the fifth year, the plumage darkens.[4][23][7]
A profile view of a wedge-tailed eagle at Captains Flat. The species is the largest Australian bird of prey and one of the largest eagles in the world.
The female wedge-tailed eagle is one of the world's largest eagles.[25] Its nearest rival in Australia for size is some 15 per cent smaller linearly and 25 per cent lighter in weight.[8] As is typical in birds of prey, the female is larger than the male. Although a few individual females are larger by only a small amount, they average up to 33 per cent larger.[4] A full-grown female weighs between 3.0 and 5.8 kg (6.6 and 12.8 lb), while the smaller males weigh 2 to 4 kg (4.4 to 8.8 lb).[7][6]
Total length varies between 81 and 106 cm (32 and 42 in) and the
wingspan typically is between 182 and 232 cm (6 ft 0 in and 7 ft 7 in).[4][6] In 1930, the average weight and wingspans of 43 birds were 3.4 kg (7.5 lb) and 204.3 cm (6 ft 8 in).[10][26] The same average figures for a survey of 126 eagles in 1932 were 3.63 kg (8.0 lb) and 226 cm (7 ft 5 in), respectively.[27]
According to one guide, the mean body mass of male wedge-tailed
eagles is 2.5 kg (5.5 lb) while that of females is listed as 4.7 kg
(10 lb), which, if accurate, is one of the most extreme examples of size
sexual dimorphism known in any bird of prey.[28][29]
However, another sample showed far less stark size differences, with 29
males weighing an average of 3.13 kg (6.9 lb) and 29 females an average
of 3.8 kg (8.4 lb).[30] In the same sample, from the Nullarbor Plain,
males averaged wingspan of 193 cm (6 ft 4 in) (sample of 26) and body
length of 85.2 cm (33.5 in) (sample 5) while females had an average
wingspan of 209 cm (6 ft 10 in) (sample 23) and body length of 92.1 cm
(36.3 in).[31]
However, the Nullarbor Plain eagles appear slightly smaller than
wedge-tailed eagle sizes from other surveys, based on body mass and wing
chord sizes.[9][31][32] An average length for males of 91 cm (36 in) and 100 cm (39 in) was described for wedge-tailed eagles in Queensland.[33] Another source claimed an average male weight of 3.2 kg (7.1 lb) and average female body mass of 4.3 kg (9.5 lb).[8] Yet another book lists males as averaging 2.95 kg (6.5 lb) and females as averaging 3.97 kg (8.8 lb).[24] A sample of 10 males averaged 3.14 kg (6.9 lb) while 19 females weighed 4.18 kg (9.2 lb).[22][32] The mean body mass of males in Tasmania was 3.5 kg (7.7 lb) while that for females was 4.1 kg (9.0 lb).[7]
The largest wingspan ever verified for an eagle was for this
species. A female killed in Tasmania in 1931 had a wingspan of 284 cm
(9 ft 4 in), and another female measured barely smaller at 279 cm (9 ft
2 in).[27] Similar claims, however, have been made for the Steller's sea eagle (Haliaeetus pelagicus),
which has been said to reach or exceed 274 cm (9 ft 0 in) in wingspan.
Reported claims of wedge-tailed eagles spanning 312 cm (10 ft 3 in) and
340 cm (11 ft 2 in) were unverified and deemed to be unreliable per Guinness World Records.[27] This eagle's great length and wingspan place it among the largest eagles in the world, but its wings, at more than 65 cm (26 in), and tail,
at up to 45 cm (18 in), are unusually elongated for its body weight,
and nine or ten other eagle species regularly outweigh it.[4] It is around the third heaviest Aquila species, outsized only somewhat by the golden eagle and slightly by the Verreaux's eagle, although it only slightly exceeds the weight of the Spanish imperial eagle (Aquila adalberti).[8][34][35] Among the entire booted eagle subfamily, in addition to the two heavier Aquila, it is outsized in bulk by the martial eagle (Polemaetus bellicosus), while the also long-tailed crowned eagle (Stephanoaetus coronatus) can average of a roughly similar body mass to the wedge-tailed eagle, although the latter is marginally the heavier bird.[8][34][22]
The wedge-tailed is exceeded in body mass by only a few eagles, especially the Steller's sea eagle and harpy eagle (Harpia harpyja) and somewhat by the Philippine eagle (Pithecophaga jefferyi), the white-tailed eagle (Haliaeetus albicilla) and the bald eagle (Haliaeetus leucocephalus).[34][25]
However, it rivals the Steller's and harpy eagles and is known to be
exceeded only by the Philippine eagle in total length. The wedge-tailed
eagle's wingspan is the largest of any Aquila, and is exceeded
amongst all eagles probably only by the white-tailed and Steller's sea
eagles in average spread though its average (not maximum) wingspan is
rivaled by that of the martial eagle.[27][25][36][37] Among standard measurements, within the nominate subspecies, the wing chord of males may range from 553 to 667 mm (21.8 to 26.3 in) while that of the female is from 600 to 703 mm (23.6 to 27.7 in).[4]
Captive specimen at Symbio Wildlife Park, New South Wales, Australia. Note the massive, pale bill and cere.
In Tasmania, the wing chord measured from 594 to 664 mm (23.4 to
26.1 in) in males and 620 to 711 mm (24.4 to 28.0 in) in females.[20] In Nullarbor Plain, males averaged 587 mm (23.1 in) in wing chord while females averaged 638 mm (25.1 in).[31] Other Australian wedge-tailed eagles averaged 611 mm (24.1 in) in wing chord among males and 650 mm (26 in) among females.[32] In Tasmania, the wing chord averaged 618.8 mm (24.36 in) in males and 661.3 mm (26.04 in) in females.[20] The extreme tail length, slightly to greatly exceeding that of other Aquila,
is in males from 352 to 479 mm (13.9 to 18.9 in), averaging 370 mm
(15 in) in the Nullarbor eagles and 421.2 mm (16.58 in) in Tasmania, and
from 376 to 536 mm (14.8 to 21.1 in), averaging 410 mm (16 in) in
Nullarbor and 448.9 mm (17.67 in) in Tasmania. Although they only
slightly exceed in tail length the heavier two Aquila and crowned eagles and they can rival the tail lengths of the Philippine and the Harpiinae eagles, Tasmanian wedge-tailed eagles are quite likely to be the longest-tailed of all modern eagles.[4][8][20][31]
The length of the tarsus may be from 99 to 139.9 mm (3.90 to 5.51 in).[4][20] The tarsus of 7 males averaged 104.3 mm (4.11 in) while that of 7 females averaged 111.1 mm (4.37 in).[22] In terms of bill
measurements, the exposed culmen may range from 37.5 to 61.6 mm (1.48
to 2.43 in) in males and 46.3 to 65.1 mm (1.82 to 2.56 in) in females
while total bill length (from the gape)
is from 55 to 67 mm (2.2 to 2.6 in) and 58.2 to 73 mm (2.29 to
2.87 in), in the sexes respectively. It is likely to be the largest
billed Aquila, a bit ahead of the imperial eagles and the Verreaux's eagle, behind only the larger Haliaeetus and Philippine eagles amongst all eagles.[20][31][32]
In Tasmania, culmen lengths averaged 48.4 mm (1.91 in) in males and
51.4 mm (2.02 in) in females while the total length of the bill averaged
59.4 mm (2.34 in) and 63.2 mm (2.49 in).[20]
The hallux claw,
the enlarged rear talon on the hind toe, is slightly smaller than that
of a golden or Verreaux's eagle, even proportionately, but is extremely
sharp.[22]
According to one study, wedge-tailed eagles had a hallux claw of
38.9 mm (1.53 in), ranging from 38.1 to 41 mm (1.50 to 1.61 in), in
males 44.7 mm (1.76 in), ranging from 40.6 to 48.1 mm (1.60 to 1.89 in)
in a sample of 10, in females.[22]
Another source listed the hallux claw of mainland Australian eagles as
averaging 41 mm (1.6 in) in males and 46 mm (1.8 in) in females.[8]
Meanwhile, in Tasmanian eagles, the hallux claw averaged 45.2 mm
(1.78 in), ranging from 38.6 to 48.8 mm (1.52 to 1.92 in) in males while
in females, the hallux claw averaged 49.9 mm (1.96 in), ranging from
45.5 to 55.6 mm (1.79 to 2.19 in).[20] In terms of osteological
structure and size, the wedge-tailed eagle is said to be proportional
to other eagles, being notably smaller and less robust than the heaviest
eagles, such as Steller's and harpies, but fairly similar in osteology,
in both structure and proportions, to the golden eagle.[38]
A juvenile in the Northern Territory, notable for paler areas on the wing coverts and nape as well as for a browner overall hue.
Their unique combination of large size, lanky build, long,
diamond-shaped tail (though can be round-ended when both central
feathers are moulted together), mainly black or rather dark plumage, and
long wings seen when soaring or gliding make all ages of the
wedge-tailed eagle fairly unmistakable in the majority of their range.[4][9] The only main confusion species is often the black-breasted kite (Hamirostra melanosternon),
which is surprisingly similar in colouring but is much smaller with a
relatively short, squared tail and extensive clear white windows
covering a good part of their wings.[4][7] Juveniles of the white-bellied sea eagle (Haliaeetus leucogaster),
at times mentioned as potentially confusable with a young wedge-tailed
eagle, are much paler below with a rather different flight pattern: a
short pale tail, bare legs, shorter, broader wings held in stiff dihedral.[4][8]
In New Guinea, the Gurney's eagle
is more similar than those species in form and build but the Gurney's
is somewhat smaller and more compact than the wedge-tailed eagle with
rich yellow feet, a rather shorter rounded or faintly wedge-tipped tail,
shorter and relatively broader wings (in adaptation to more
forest-living).[4][7][39] Furthermore, the Gurney's eagle has a much paler immature plumage.[4][39]
Although usually considered an island endemic, the Gurney's eagle is
possibly capable of marine dispersals, as is the wedge-tailed eagle,
that may lead to them to turn up in the forests of northern Australia
and historical reports show that a rare vagrant of the species may
indeed appear there.[8][40] The Papuan eagle (Harpyopsis novaeguineae),
the only other island raptor in New Guinea that approaches the
wedge-tailed in size, is a highly distinct and forest-restricted
species, being much paler, particularly below, with long, bare legs and
different proportions, more like a giant Accipiter with short rounded wings, a long, somewhat rounded tipped tail, and a large, rounded head.[4][39]
Wedge-tailed eagles are not well known for its vocalization nor are they often heard.[4][10] They may be silent for long stretches of time, possibly months, at least outside of breeding season.[4]
When vocalizations have been documented, it usually only near the nest
and in aerial display, and can be hard to hear unless at close range.[4] The commonest calls for wedge-tailed eagles are high, rather thin whistles, sometimes transcribed as I-see, I-see followed by a short descending see-tya.[4][7][10]
Also documented during the breeding season are various other whistles,
yelps and squeals and an often rolling series. Characteristically, all
their calls are surprisingly weak, though the main call is sometimes
considered to have a "melancholy" quality.[4][10] The opinion on their call is not dissimilar to the golden eagle, whose voice is similarly considered unimpressive.[10][41] Female calls in wedge-tailed eagles are similar but are generally lower and harsher than males.[4]
A wedge-tailed eagle seen over well-wooded environments around Kergunyah.
Wedge-tailed
eagles favor varied environments, mostly those with some leafy trees
and rocky areas, but can be seen in a great variety of habitats.
The wedge-tailed eagle lives in an extremely wide range of habitats.[4][15]
Although range is restricted relative to the golden eagle, it likely
occupies a wider range of habitat types than likely any other Aquila
eagle, and may outrival any booted eagle species in their use of
diverse habitats, being somewhat more akin to habitat generalist raptors
such as Buteo buzzards.[8][52][53] Assorted habitats known to host wedge-tailed eagles includes open woodland, savanna, heathland, grasslands, desert edge and semi-desert, subalpine forests, montane grasslands and mountain peaks, not-too-dense tropical rainforests, monsoon forests, dwarf conifer forests, some wetlands as well as regularly forays to coastal areas, though normally along the coasts they occur around plains somewhat away from the water.[4][7][8][9][15][excessive citations]
Favored habitat tends to be remote or rough country, at least partially
wooded and not uncommonly varied with some rocky spots as well as in shrubland. Wedge-tailed eagles seem to prefer some dead trees to be present.[7][8][6][15] They may occur around Eucalyptus woodland quite regularly, as well as Acacia woodland and mixed woodlands of Casuarina cristata-Flindersia maculosa-Callitriscypresses and also stands of Casuarina cunninghamiana.[7][54][55][56] A strong preference was detected for C. cunninghamiana alternatively with several Eucalyptus species was detected in the Australian Capital Territory, sloping ground allowing good access and access to tall, mature trees being paramount to the eagles in the study.[57] Quite often they will be seen soaring over hills, mountains or escarpments as well as over flat plains, especially spinex grassland.[6][15] Dense forest is typically avoided with glades and edge often sought out in forested areas. While they do occur in rich riparian woodlands, it is with relative scarcity despite this being where many other raptors of the nation concentrate.[7][6][58][59] In the deserts of the Lake Eyre basin, they are often seen in gibber plains along treedwatercourses and drainage basins, here often concentrated around Eucalyptus in stony creek beds.[7][60] In the sandy desert areas of Western Australia, wedge-tailed eagles were once reasonably common but have largely vacated the region after the macropod prey they live off of there were all but hunted to extinction.[61]
Wedge-tailed eagles commonly occur from sea level up to about 2,000 m
(6,600 ft) with seemingly no preference based on altitudinal level.[4] A fairly pronounced liking for mountainous localities such as plateaus has been detected in a few studies of wedge-tailed eagle.[58][62] One of the few habitat types considered to be strongly avoided by wedge-tailed eagles are areas intensively settled or cultivated areas.[4][9][24]
A slightly fading tendency to avoid human areas has been detected,
perhaps as persecution rates have gone far down, and the wedge-tailed
eagle may be seen near towns and villages in exurban and even suburban areas largely within bushland. However, the species is seldom seen other than as a flyover in more developed towns and cities.[7][8] Additionally, it is not uncommon to see these eagles in man-made spots such as pasture areas, forestry clearings, and rolling farmland areas.[8]
Taking off from its perch, the long legs of this adult female are clearly visible.
This impressive bird of prey spends much of the day perching in
trees, on rocks as well as similar exposed lookout sites such as cliffs
from which it has a good view of its surroundings. Alternatively, they
may sit on the ground for long periods of time or watch from a lower
point, such as on termite mounds or anthills.[4] Now and then, it takes off from its perch to fly low over its territory.[8] Especially whilst not breeding, wedge-tailed eagles spends a considerable amount of the day on the wing.[4][8]
Wedge-tailed eagles are highly aerial, soaring for hours on end without
wingbeat and seemingly without effort, regularly reaching 1,800 m
(5,900 ft) and sometimes considerably higher. The purpose of soaring has
received little specific study in wedge-tailed eagles, but it is
likely, as in other accipitrids, in large part for surveying the
territory and advertising their presence to other eagles.[53][63]
During the intense heat of the middle part of the day, it often soars
high in the air, circling up on the thermal currents that rise from the
ground below.[8] Often when on the wing, it is scarcely visible to the human's naked eye.[4] Their keen eyesight extends into ultraviolet bands. With a visual perception some three times more acute than those of humans, one of the largest pecten oculi of any bird and an eye roughly as big as a small human's, they may be one of the most sharp-eyed birds in the world.[8][64][65]
The wedge-tailed eagle is largely sedentary as expected of a raptor dwelling in the subtropics, although they also dwell in the tropics (far northern Australia and New Guinea) as well as in the temperate zone (Tasmania).[4][10]
However, juveniles of the species can be quite dispersive. In some
cases, they have moved to a recorded distance of some 836 to 868 km (519
to 539 mi). These extreme movements have been completed within 7 to 8
months after dispersal.[8][9] More typically they move no farther than 200 km (120 mi) or so.[24] The adult eagles can also be nomadic, though only in circumstances such as drought conditions. In turn this explains the species presence in places they don't breed, even adults.[4][8][9] In addition to moving for drought in arid zone, also moves in highest part of New South Wales, e.g. the Snowy Mountains, the species often apparently vacates snow-covered alpine zone in winter.[66]
The small New Guinea population is apparently indistinguishable from
the mainland race and so possibly result of recent colonization,
although no records exist of migrating wedge-tailed eagles islands past
the Torres Straits.[4]
However, it can be projected from its presence in various offshore
islands its capacity for crossing straits ranging up to as far as 50 to
100 km (31 to 62 mi) apart.[8] One post dispersal young eagle was observed to distribute from Kangaroo Island to the mainland, possibly a regular occurrence.[67] Due to their tendency for wandering, some authors class the wedge-tailed eagles as a “partial or irruptive migrant”.
However, while they are arguably irruptive, it does not fit the mould
of a true migrant well since under normal circumstances adults are
rather sedentary unless environmental changes force them to move.[15][68]
The wedge-tailed eagle is the only bird that has a reputation for not infrequently attacking hang gliders and paragliders,
although other eagles including the golden eagle have also been
recorded to behave thusly. Based on the response the eagles show to the
gliders, they presumably are defending their territory and treating the
perceived intruder like another eagle. Cases are recorded of the birds
damaging the fabric of these gliders with their talons as well as some
other parts of the gliding apparatus, but not the humans themselves, has
been reported.[8][69] They have also been reported to attack and destroy unmanned aerial vehicles used for mining survey operations in Australia.[70]
The presence of a wedge-tailed eagle often causes panic among smaller birds and, as a result, aggressive species such as magpies (one of the most vulnerable types of passerine to eagle attacks), butcherbirds, wagtails, monarch flycatchers, lapwings, and miners as well as smaller birds of prey, including both accipitrids and falcons, any of which may aggressively mob
eagles (see video). Multiple species may join the kerfuffle and mob
them, especially while the eagles perched, often engaging in noisy
calling, presumably meant to disorient the predator, and occasionally in
physical attacks against the eagle, typically focused where the big,
relatively lumbering eagles could not grasp the attacking birds. The
wedge-tailed eagle usually does not engage its tormentors but sometimes
rolls in the air to present talons whether perched or not.[8][71][72]
Sometimes wedge-tailed eagles appear to fight but this and other
behaviours, especially between young eagles, may be interpreted as
playful.[8][9]
Some such behaviours have included fetching sticks tossed by others,
athletic flipping between juvenile eagles and even playing games with dogs, via floating above them until the dogs bark or leap then floating up until the dog settles and then repeating the “game”.[8]
Flocking behaviour has been noted, similar to that of vultures (Cathartidae and Accipitridae) in other countries, when carrion is available.[73]
Duration: 38 seconds.0:38
In flight, 'mobbed' by Australian magpie, Dayboro, SE Queensland
Duration: 41 seconds.0:41
Samsonvale, SE Queensland, Australia
The wedge-tailed eagle is one of the world's most powerful avian predators.[7][8] Due to its formidable and dominating nature, it is sometimes nicknamed “King of Birds”, along with golden eagles.[74]
Prey is usually grabbed via a pounce or snatch during a gliding flight
or a tail-chase from low quartering or transect flights.[4][7][59][75] Prey is not infrequently spotted from a soaring flight and they may undertake a long, slanting stoop towards it.[10] They may be able to spot prey from farther than a kilometre given their keen vision.[8] Its typical hunting style is not all-together dissimilar from that of golden or Verreaux's eagles.[53] Occasionally, a wedge-tailed eagle still hunts from a perch.[4] Unsuccessful hunts typically exceed in number successful ones.[10]
Hunting habitat can be highly variably and can manage to capture
prey in both open country and quite thick woodland or forest, though
typically require an open understory in the latter.[10] Almost all its prey is taken on the ground but to a lesser extent it may be taken from the tree canopy.[4] They've been known to take birds such as currawongs and cockatoos by coming around them by surprise around a tree or by darting out in flight at close range for a brief tail-chase.[8][76] Sometimes, an eagle may pull brushtail possums and other mammals from tree cavities, as well as young birds from a nest.[4][77][78] They've been known to follow wildfires to search for fleeing animals or alternately tractors and other farm equipment for the same purpose.[8] Wedge-tailed eagles occasionally pirate food from other predators.[4][8] An eagle of the species can carry prey of at least 5 kg (11 lb).[4][7]
Large animals may be attacked by pairs or, occasionally, by groups acting cooperatively. One record shows 15 wedge-tailed eagles hunting kangaroos, two actively chasing at a time, then repeatedly being replaced by two more from the circling group overhead.[4][8][79][80]
Regardless of prey size and season, tandem hunts, mainly by breeding
adult pairs or sometimes loosely associated young eagles, are not
uncommon.[7][9] Of 89 observed hunts in Central Australia around one-third were cooperative ones.[7]
As in other tandem hunting raptors, one eagle typically lies in wait
generally unseen while the other eagle distracts and drives the prey
towards it.[7][81] When hunting domesticated prey, they've been seen to land near livestock mothers to intimidate them and separate their young, so they can attack the latter.[7] Sometimes, wedge-tailed eagles may use fences to limit a prey's escape routes.[7]
In some cases, these eagles will attempt to force large prey such as
kangaroos and dingos to fall off steep hillsides and injure themselves.[8]
At times, wedge-tailed eagles appear to hunt at earliest light or late twilight in order to come upon nocturnal prey such as hare-wallaby and bettongs.[8] These eagles have been seen removing rabbits from traps and eating carrion in bright moonlight as well.[8] At times, remarkably, wedge-tailed eagles have been covering large prey with vegetation, apparently to cache food too heavy to carry.[82] Carrion is a major diet item, also; wedge-tails can spot the activity of ravens
around a carcass from a great distance, and glide down to appropriate
it. Carrion consumption is recorded in all season and contexts, although
generally non-breeding birds are more likely to scavenge
and young wedge-tailed eagles, even more so shortly post-dispersal, are
thought to be far more likely to scavenge on carrion than adults
generally. Wedge-tailed eagles are often seen by the roadside in rural
Australia, feeding on animals that have been killed in collisions
with vehicles. The importance of carrion relative to live prey has not
been greatly studied but away from human development, especially roads,
carrion is less likely to be encountered and eagles of all ages must
presumably hunt to survive. In general, Australian accipitrids of many species not infrequently come to carrion and they along with large passerines like Corvus species and currawongs probably fulfill the niche that vultures do in other continents to some extent, albeit with considerably less specialization.[4][7][8][9]
Aggregations of wedge-tailed eagles may occur not infrequently at
large carcasses, with up to 5–12 eagles or sometimes 20 gathering.[4]
A wedge-tailed eagle can gorge up to 1 to 1.5 kg (2.2 to 3.3 lb) at a
sitting and, when fulfilled, can lasts for an unusual amount of time,
for up to weeks or even a month, before needing to hunt again,
apparently due to the warmth of the environment.[7] After feeding they may disgorge a relatively small pellet, 29 to 98 mm (1.1 to 3.9 in) long by 20 to 50 mm (0.79 to 1.97 in) wide and weighing some 8.8 g (0.31 oz).[7] Usually the diet is determined from a combination of reviewing these pellets along with loose prey remains.[83]
A diversity of birds may be taken with medium-sized, common birds such as galahs taken relatively frequently due to the conspicuousness.
The wedge-tailed eagle is a dietary generalist, opportunistically capturing a wide range of prey species.[7][9]
Its prey spectrum is quite broad, with well over 200 prey species
documented to be taken and even this includes very few prey only from
secondary accounts from Tasmania and New Guinea.[7][8][9][84] The wedge-tailed eagles tends to prefer smallish to fairly large mammals as prey.[4][7] However, they not infrequently take ample numbers of both birds and reptiles, along with scarcely other prey taxon.[7][9][84]
Out of 21 accrued dietary studies, 61.3% of prey items by number in the
foods during nesting efforts were mammals, 21.6% were birds, 13.2% were
reptiles, 2.1% by invertebrates, principally insects, 1.5% by fish, and almost no amphibians by number.[7] Meanwhile, out of the 21, 13 studies calculated estimated biomass, and found that just shy of 90% of prey biomass was made of by mammals, 6.2% by birds and 3.4% by reptiles.[7] Out of the Aquila genus, it is one of a few generalist species, however the wedge-tailed eagle is the Aquila most likely to typically attack the largest prey.[53]
Generally, this species prefers to attack birds and reptiles weighing
over 100 g (3.5 oz) and mammals weighing over 500 g (1.1 lb), although
prey taken at times has varied from a few grams to more than sixteen
times the weight of an individual eagle.[8][9]
A comparison estimate posited that around 2% of wedge-tailed
eagle prey weighs less than 63 g (2.2 oz), 4% of their prey weighs 63 to
125 g (2.2 to 4.4 oz), 7% of their prey weighs 125 to 250 g (4.4 to
8.8 oz), 10% weighs 250 to 500 g (8.8 to 17.6 oz), 20% weighs 500 to
1,000 g (1.1 to 2.2 lb), 25% weighs 1,000 to 2,000 g (2.2 to 4.4 lb),
18% weighs 2,000 to 4,000 g (4.4 to 8.8 lb) and 14% weighs over 4,000 g
(8.8 lb).[53]
Projected from this comparison, the mean prey size for wedge-tailed
eagles is estimated at 1,750 g (3.86 lb), similar but just slightly
ahead of the Verreaux's eagle and some 14% ahead of the golden eagle
global mean prey size.[53] Further studies estimated mean prey weight, showing the mean prey weigh in the Canberra-Australian Capital Territory
region in three different studies was estimated to be 1,298 g
(2.862 lb), 2,131 g (4.698 lb) and 2,890 g (6.37 lb), changing likely
due to the shifting significances of leporids and larger macropods.[84][85][86] In a small study from Armidale, New South Wales, it was estimated that mean prey weight was 1,309 g (2.886 lb).[87]
It only ranks behind the crowned eagle and harpy eagle and rivals the
martial eagle as the eagle likely to attack the largest prey on average.[88][89][90]
While the introduction of invasive species
to Australia has been generally having a negative to devastating effect
on native animals and ecosystems, the wedge-tailed eagle is one of a
few native species to largely benefit from these introductions.[7][91] This is especially due to the introduction of the European rabbit,
which were deliberately introduced repeatedly (abortively in 1859 and
then via a concerted effect from 1937 to 1950), largely so the wealthy
could hunt them.[92] The wedge-tailed eagles quickly took to the rabbits as prey along with another introduced leporid, the European hare (Lepus europaeus).[7][8][9] In almost every part of Australia, these eagles take rabbits in some numbers and they usually constitute the bulk of the prey species in most, if not all, Australian food studies.[84][85] In some dietary studies rabbits have accounted for up to 89.2% of the diet by number and 86% by biomass, as in Bacchus Marsh, however they more typically range from 16% to 49% of the diet by number in various studies.[85][93][94] One Canberra study found that 98.5% of the rabbits taken were adults.[84]
In the largest study near Canberra, over 5.5 years, 19.3% of the diet
of wedge-tailed eagles was rabbits (12.7% of prey biomass) among 1421
prey items, so the eagles took a total of some 275 rabbits in the 11 to
17 studied territories of the area.[84] A study estimated that mean weight of wild rabbits in Australia was 1.4 kg (3.1 lb), lower than estimated in the past.[58]
However, other studies estimated the mean weights of rabbits taken by
wedge-tailed eagles as variously from 1.5 to 2.2 kg (3.3 to 4.9 lb) or
“usually over 1.65 kg (3.6 lb)”, infrequently reported to 2.47 kg
(5.4 lb), size of the rabbits being perhaps limited the poorly-suited
soil and environs of the Australian wilderness.[85][93][58][95]
Meanwhile, the European hare is neither as widely established nor as
prolifically taken as rabbits by wedge-tailed eagles but are by no means
neglected and a substantial meal.[84][85]
With a mean body mass of 4 kg (8.8 lb), hares have been as much as
nearly 10% of the local diet and up to 14% of prey biomass in studies.[84][85][93]Rabbit haemorrhagic disease
was deliberately introduced to control the population of rabbits
subsequent to 1995, followed more effectively by introduction of myxoma virus to limit the damage rabbits have inflicted on native vegetation and resultingly have competed native mammals like wallabies out of parts of their range.[92][96] Ultimately, the rabbit population may have more than halved and locally have been some 90% reduced.[8][92]
As a matter of consensus, the wedge-tailed eagles do not appear to be
adversely affected in major ways by the biological control of rabbits
since they can revert to primarily taking native prey species quite
readily.[97] In the region of Broken Hill, White Cliffs and Cunnamulla, rabbits have gone down from accounting for 56–69% of the diet to 16–31% of it.[58]
Furthermore, wedge-tailed eagles have been known to successfully
maintain population in the absence of any rabbits in a few areas.[7][98]
Wedge-tailed eagles are disproportionately likely to be seen eating carrion of roadkills (killed by cars). In the wild they often capture live prey. They often tolerate sharing carrion with other scavengers, such as ravens, despite them being prey species as well.
Much more controversial at one time than hunting introduced rabbits
and hares is the wedge-tailed eagle's occasional tendency to feed on and
sometimes kill domesticated livestock animals.[99][100]
The predation of wedge-tailed eagles on young farm animals has been the
primary historic driver for the persecution of the species.[101] However, in no known study have domestic livestock been known to be primary prey.[7][8] The closest association with them was in northwestern Queensland where lambs (Ovis aries) made up 32.7% of prey in pellets and 17.1% in remains, accounting for 15–21% of the prey biomass, while juvenile pigs (Sus scrofa domesticus) made up 7.3% of pellet remains and 22% of the biomass.[100]
Although it can be highly difficult, attempts have been made at parsing
out whether the eagles had indeed killed the lambs rather than just
lifting or dismantling them after finding them dead, as this eagle quite
readily comes to carrion. The findings were that of 29 diagnosable lamb
deaths in northwest Queensland, only 34.5% were due to eagle attacks.[9][100]
The wedge-tailed eagle is at times capable of taking very substantial
livestock animals, lambs taken have been estimated to weigh a mean of
3.5 kg (7.7 lb) or up to 15 kg (33 lb) while fully grown sheep weighing
some 40 to 50 kg (88 to 110 lb) are infrequently vulnerable, presumably
in large part to hunting pairs of eagles.[85][86][58][100]
In the largest study of the Canberra area, 82.5% of diagnosable sheep
specimens were adults but probably were by and large scavenged.[84] Meanwhile, young pigs included in the diet were estimated to weigh around 14 kg (31 lb), and sometimes feral piglets are included in the diet.[95][100]
When attacking lambs, the wedge-tailed eagles are apparently capable of
driving their talons into the skull of the victim, although more
typically they land along the back and grip the lamb along the spine
until it weakens and collapses while flapping the wings for balance.[99][100]
This species will also land between a ewe or female pig and their
respective lambs or piglets in order to separate the latter for attack.[7] Wedge-tailed eagles are also known to at times prey on another animal introduced for human hunting purposes, the red fox (Vulpes vulpes),
which can form up to about 4% of an eagle's breeding diet and 5% of the
biomass, weighing up to 9 kg (20 lb). In Canberra, about 59% of the
foxes found in the diet were adults.[84][86][102][103] Additionally, feral cats, mainly juveniles, can be part of their prey.[102][104]
Presumably, the primary native prey of wedge-tailed eagles is marsupials, particularly macropods, which is also in accord with studies involving places where rabbits have declined or never occurred.[7][98][105]
Many wallabies, kangaroos, and associated animals are included in the
diet, with over 50 marsupials known to be in the species’ prey spectrum.
When selecting marsupials, wedge-tailed eagles tend to ignore smaller
species and focus on larger-sized ones.[7][9][84][85] However, they generally most often take alive the young, small and sickly of large macropod marsupials.[85][28]
Findings were that juvenile macropods were taken out of proportion to
their numbers in the environment, unlike rabbits which were taken
roughly in proportion to their abundance.[55]
In recent times, they have been known to eat marsupials, such as
kangaroos killed by cars. There is little evidence that macropods
delivered to nests are usually roadkills or from carrion, but the source
of prey is difficult to determine because, to minimize disturbance,
examinations are usually done after breeding is complete. As well, the
attendance at carrion by wedge-tailed eagles is disproportionately done
by juvenile eagles. In one study of roadkills in Australia, the species
ranked around fourth in frequency and capacity for carcass breakdown of
scavengers at roadkills, behind feral pigs, red foxes, and ravens.[59][106][107]
A video surveillance study at the nest determined that seemingly
freshly killed, albeit usually quite young macropods were delivered to
nests near Broken Hill.[55] As much as 20% to 30% of the diet can be made up of by macropods.[7] Large and prominent species are known including the grey kangaroos and the red kangaroo (Osphranter rufus). Generally, juveniles are targeted of these large species with eastern grey kangaroos (Macropus giganteus)
estimated to weigh 17.2 to 20 kg (38 to 44 lb) when taken by
wedge-tailed eagles, in Australian Capital Territory and New South
Wales, while the weight of young western grey kangaroos (Macropus fuliginosus) was said to be 3.8 kg (8.4 lb) in one study in Western Australia.[85][86][28]
The estimated weight of juvenile red kangaroos taken was 9 kg (20 lb)
in northwestern Queensland where they were the primary prey species
ahead of lambs.[100] However, wedge-tailed eagles do not shy away from attacking large, adult macropods.[8][9] Similarly large adult macropods killed by these eagles can include common wallaroos (Osphranter robustus) (mean adult weight around 25 kg (55 lb)), antilopine kangaroo (Osphranter antilopinus) (mean adult weight around 30 kg (66 lb)), agile wallaby (Notamacropus agilis) (median adult weight around 16.8 kg (37 lb)), black-striped wallaby (Notamacropus dorsalis) (median adult weight around 13 kg (29 lb)), red-necked wallaby (Notamacropus rufogriseus) estimated to weigh around 16 kg (35 lb) when taken, swamp wallaby (Wallabia bicolor) (mean adult weight around 15 kg (33 lb)), and even red kangaroo adults.[7][85][86][100][108][excessive citations] They've been recorded attacking eastern grey kangaroos weighing over 35 kg (77 lb).[8] In extreme cases, wedge-tailed eagles have killed kangaroos weighing approximately 60 kg (130 lb).[8]
In one case, a huge male eastern grey kangaroo, estimated to stand
1.7 m (5 ft 7 in) was successfully dispatched by a pair of wedge-tailed
eagles.[109]
Furthermore, an adult female western grey kangaroo was witnessed to be
killed “in a few minutes” by a hunting pair of wedge-tailed eagles, and
the eagles are considered a serious predator of the western grey.[110]
In some unusual cases, wedge-tailed eagle hunting parties can form
whilst hunting red kangaroos, sometimes including up to 15 eagles (more
loose, opportunistic aggregations than well-organized groups), but usually only a pair is sufficient to kill such prey.[9][109]
Normally, the eagles repeatedly attack the kangaroo, sinking their
talons into the back or nape and then fly up, when the second eagle
starts doing the same. In some cases, as many as 123 attacks have been
carried out against large kangaroos before they succumb.[9][109]
When attacking joeys, eagles may, in some cases, have intentionally
caused a mother kangaroo to dislodge a joey from the pouch to capture
and fly off with it.[109]
With relative infrequency, other classes of mammals, beyond
leporids and marsupials, may be taken opportunistically by wedge-tailed
eagles.[7] At least two species each of flying foxes and wattled bats are included in the prey spectrum.[7][9][132][133] Occasionally, an eagle may take a monotreme including both the platypus (Ornithorhynchus anatinus) and the short-beaked echidna (Tachyglossus aculeatus).[84][134] Several species of rat are readily taken and even the house mouse (Mus musculus), likely the smallest mammalian prey known for wedge-tailed eagles at around 20 g (0.71 oz) in weight.[84][93][102] Although rare, a dingo (Canis familiaris) may be taken by a wedge-tailed eagle at times, mostly pups, or carrion but sometimes a pair of eagles can kill adults too.[8][135][136] Beyond sheep, pigs, and infrequently young goats (Capra hircus), other ungulate prey, entirely introduced by man into the Australasian region, is eaten exclusively as carrion so far as is known, including cattle (Bos taurus - despite claims that eagles have killed young calves, which is possible, they have only ever been witnessed feeding on afterbirths and not harming calves), Javan rusa (Rusa timorensis) in New Guinea, sambar deer (Rusa unicolor) in northern Victoria and water buffalo (Bubalus bubalis) in the Northern Territory.[84][137][138][139][140][excessive citations]
In one instance, a young girl was apparently subject to a brief attack
by a wedge-tailed eagle, in what was likely an attempted act of
predation, near her rural home but the attack was abandoned by the
eagle. It has been noted that some different species of large eagles are
thought to occasionally attack children as prey though, among extant
species, only the crowned eagle and martial eagle, both in Africa, are thought to have successfully carried out rare acts of predation on human children.[8][141]
A wedge-tailed eagle feeding.
Birds take a clearly secondary position to mammals when importance
and especially prey weight are concerned; however, the wedge-tailed
eagle shows some fondness for avian prey.[7][8] With more than 100 prey species included in the prey spectrum, birds are the most diverse class of prey taken by these eagles.[7][84][85][102] Generally, the predation of birds seems to be highly opportunistic and no one type of bird reliably dominates the eagle's diet.[7][102]
However certain species, probably due to their commonality in eagle
territories and perhaps vulnerability through their own behaviour that
seem to be taken most often.[7][84] These consist of Corvus species, especially little (Corvus mellori) and Australian ravens (Corvus coronoides), weighing a mean between species of around 580 to 590 g (1.28 to 1.30 lb) when taken, 300 to 329 g (10.6 to 11.6 oz) Australian magpies (Gymnorhina tibicen), 800 to 820 g (1.76 to 1.81 lb) Australian wood duck (Chenonetta jubata), 310 to 335 g (10.9 to 11.8 oz) galah (Eolophus roseicapilla), larger cockatoos and smaller parakeets and parrots.[84][85][86][102][142][143][excessive citations] On Kangaroo Island, Australian and little ravens together constituted 19% of the diet.[98] In Canberra, fairly prominent numbers of magpies, wood ducks, galahs and eastern (Platycercus eximius) and crimson rosellas (Platycercus elegans) are known to be taken, these collectively forming up to about 25% of the diet by number.[85]
In the Perth region, birds were taken amply, especially the Australian
raven at 12.6% of prey remains and 4.7% of the biomass, with birds
constituting just shy of 25% of the diet.[28] Elsewhere in Western Australia, a similar percentage of the diet is made up of by birds, mostly the same species with some number of Australian ringnecks (Barnardius zonarius) and Baudin's black cockatoos (Zanda baudinii) as well.[102][144] Peculiarly, one study found that among a large sample of 1826 prey items in the Northern Territory that the most often identified prey species was the tiny budgerigar (Melopsittacus undulatus), at 29 g (1.0 oz) one of the smallest avian prey species for this eagle.[34][59] In a single study from the Fleurieu Peninsula, birds were the majority of prey for wedge-tailed eagles, at 62.5%, mostly Corvus followed by wood duck, galah and magpies.[142] Other assorted avian prey include several species of waterfowl, including several ducks as well as swans and geese, and a fairly strong frequency of attacks on large rails, such as swamphens, moorhens, native-hens and coots.[7][28][59][102][139][145][146][excessive citations]
When selecting reptiles as prey, wedge-tailed eagles by far are most likely to pursue lizards.
The range of lizards they may prey upon is highly diverse in size and
nature, with somewhere between 20 and 30 species known in the prey
spectrum.[7][9][157] The most preferred reptilian prey by far is bearded dragons.[7][9]
Despite the small size of this prey relative to most mammalian prey,
they can be key to survivorship in more arid vicinities such as central
and western Australia where there is less diverse prey to pick from.[9][55] In video monitored prey deliveries at Fowlers Gap Arid Zone Research Station, central bearded dragons (Pogona vitticeps)
dominated the prey composition, making up 68.2% of 110 prey deliveries
and the only known instance of reptiles forming the bulk of wedge-tailed
eagle diet.[55]
A different study from prey remains and pellets found the central
bearded dragon to comprise 28.6% of the diet among 192 prey items.[56] In south-central Queensland, the bearded dragon was the leading prey species by number, making up 26.9% of 729 prey items.[95] In northeastern New South Wales, the eastern bearded dragon (Pogona barbata) was the second most numerous prey species behind the rabbit, at 16.6% of the diet.[121]
The bearded dragons when taken by wedge-tailed eagles have had an
estimated body mass ranging from 80 to 320 g (2.8 to 11.3 oz).[28][100] They also prey on jacky dragons.[158] Larger lizards are readily taken as well given the opportunity. Skinks are occasional supplemental prey, common blue-tongued skink (Tiliqua scincoides) at around 500 g (1.1 lb) can make up around 5% of the diet (in northeast New South Wales), while the 204 g (7.2 oz) Centralian blue-tongued skink (Tiliqua multifasciata) was quite prominent in the diet in the Northern Territory.[59][121][159] In Western Australia, 617 g (1.360 lb) shingleback skink (Tiliqua rugosa) and somewhat smaller western blue-tongued skink (Tiliqua occipitalis) collectively comprised about 7.5% of the diet.[102][159] Much bigger lizards are sometimes taken, namely monitor lizards. Around 20% of the 231 prey items of in a Western Australian study was found to be monitor lizards, mostly 2.32 kg (5.1 lb) yellow-spotted monitors (Varanus panoptes) with some 821 g (1.810 lb) sand goannas (Varanus gouldii).[120][159] Adult Rosenberg's monitors (Varanus rosenbergi), weighing around 1.1 kg (2.4 lb) can be also taken.[160] Even lace monitors (Varanus varius), which weighs 6.3 kg (14 lb) on average adults, can be a prey for this powerful eagle.[121][161][162] Contrarily, lizards down to the size of a 20 g (0.71 oz) pygmy spiny-tailed skink (Egernia depressa) and a 38 g (1.3 oz) thorny devil (Moloch horridus) may be taken.[102][120][159][163]
Beyond lizards, wedge-tailed eagles seldom seem to hunt other types of reptiles.[157] They hunt a few species of snakes, mostly venomous species, because they are prevalent in Australia. Snakes known to be included in the diet including tiger snakes (Notechis scutatus), eastern brown snake (Pseudonaja textilis), ringed brown snake (Pseudonaja modesta), bandy-bandy (Vermicella annulata), yellow-faced whipsnake (Demansia psammophis), red-bellied black snake (Pseudechis porphyriacus) and brown tree snake (Boiga irregularis).[84][102][157][164][165][excessive citations]Eastern long-necked turtle (Chelodina longicollis)
have been claimed as prey in one report although any other confirmed
cases of predation on turtles by this species are not known.[7][157] Notably, there are no reports of wedge-tailed eagles attacking pythons, despite several species being present in Australia, nor on crocodiles;
perhaps these are the only predators too formidable to be attacked, as
both of these reptiles can attain extremely large sizes.[166][167] Predation on frogs or other amphibians is almost unheard of for wedge-tailed eagles, however, based on toxicity reports in eagles, they may consume invasive cane toads (Rhinella marina) from time to time.[168] Similarly rare in the species’ diet is fish, although common carp (Cyprinus carpio) and western blue groper (Achoerodus gouldii) have been documented as prey.[84][102] Occasionally, wedge-tailed eagles may even attack insects such as Psaltoda moerens cicadas and Heteronychus arator beetles.[84]
Truly exceptional is in the Northern Territory, where a large
percentage of 1826 prey items was made up of by insects including
unidentified Orthoptera, at about 10.8% of the diet, unidentified beetles, at about 8.4%, as well as some numbers of ants.
Why and how they capture a profusion of insects locally is not clear,
and they may be often from the stomachs of other prey or even byproduct
from the captures of other prey or from the bodies of carcasses.[59]
Being the largest and most powerful Australian raptor, wedge-tailed eagles are dominant over other raptor species of the region.
The wedge-tailed eagle occupies a fairly unique niche relative to other Aquila. While primarily continental in distribution, it is distributed well apart from most related species, whereas most Aquila are distributed in Eurasia or Africa
and face considerable competition over resources, enabling certain
specializations of most species in habitat or microhabitat, morphology
and behaviours and often life history, including nesting grounds and
often foods.[10][53][169]
The wedge-tailed eagle has the ability to exploit a more catholic
variation of both prey and habitat since it exists with relatively fewer
competing species.[7][9] The most considerable potential competition comes in the two other eagles regularly distributed in Australia, the little eagle and white-bellied sea eagle.[84][85]
The little eagle has a few ecological similarities to the wedge-tailed
eagle. It is also something of a habitat generalist, although it is
found somewhat scarcely in more arid vicinity, high elevation areas and
varied semi-open forest than the wedge-tailed eagle.[7][61][84] Like the wedge-tailed eagle, the little eagle has in recent decades become a somewhat specialized predator of European rabbits.[7][84]
However, the size difference is extreme between the wedge-tailed and
little eagles, with the earlier over four times heavier than the latter,
and the little eagle as expected exploits a lower trophic level relative to its more powerful competitor.[7][84] As in other areas where booted eagles and sea eagles have abutting ranges, sometimes wedge-tailed eagles compete with white-bellied sea eagles.[85] One key difference from elsewhere where competition sometimes occurs such as the golden eagle with the white-tailed eagle (Haliaeetus albicilla) in Eurasia and the bald eagle (Haliaeetus leucocephalus)
in North America, both of which are slightly heavier than the golden,
is that the white-bellied sea eagle is the slightly smaller species than
the wedge-tailed eagle, potentially giving the latter a more pronounced
competitive edge.[7][8][170]
However, the white-bellied sea eagle clearly does not shy away from
contentious border disputes with wedge-tailed eagles and the two species
can often be seen be seen readily attacking each other, occasionally in
talon grappling and sometimes cartwheeling attacks on one another.[61][85]
However, the ecological effect of interspecific competition of the two
species is not clear. Although the wedge-tailed eagle is considered the
dominant species of the two, they clearly do not take the presence of
white-bellied sea eagles lightly and some authors feel they may avoid
nesting near them.[8][85][171] Clearly, there is ample partitioning between the wedge-tailed and white-bellied sea eagles, the latter adapted to mostly open wetlands and coasts and, while also a dietary generalist, they tend to derive most of their diet from fish, water birds and other wetland-dwelling prey, and they seldom compete directly for prey such as mammals with wedge-tailed eagles.[85][171]
Most other diurnal raptors that reside in Australia are
considerably smaller and seldom can be said to present great competition
to the wedge-tailed eagles, although some, such as swamp harriers (Circus approximans), black-breasted kites and grey goshawks (Accipiter novaehollandiae), are relatively large for their taxon and powerful predators in their own rights.[7][9] In one instance, a square-tailed kite (Lophoictinia isura) was observed to engage in an apparent territorial fight with a wedge-tailed eagle, including talon-grappling.[172]
When it comes to carrion, wedge-tailed eagles tend to dominate other
predators, especially most birds, with most kites, other assorted
raptors and some large passerine birds, mainly Corvus species and butcherbirds, coming to dead animals including roadkills.[9][173]
However, heavier terrestrial meat-eaters can hold their own at times
against wedge-tailed eagles, namely red foxes, dingos, monitor lizards
and Tasmanian devils, despite all these species sometimes turning up as
prey of these eagles as well.[7][8]
Sometimes the wedge-tailed eagle will readily rob various other raptors
of their prey, including little eagles, white-bellied sea eagles and brown falcons (Falco berigora).[7][78][174][175]
Wedge-tailed eagles will opportunistically prey on other birds of prey.
They share this aptitude with other large eagles in different parts of
the world such as golden eagles, although such acts are relatively
infrequent, it is clear that the wedge-tailed eagle is considered a
primary threat by many raptors based on witnessed attacks by eagles on
them and the mobbing behaviour of other raptors.[7][53][176] Among the other birds of prey known to occasionally fall prey to these eagles are little eagles, collared sparrowhawks (Accipiter cirrocephalus), grey goshawks, brown goshawks (Accipiter fasciatus), Pacific bazas (Aviceda subcristata), black-breasted buzzards, peregrine falcons (Falco peregrinus), Australian hobbies (Falco longipennis), black falcons (Falco subniger), brown falcons and Nankeen kestrels (Falco cenchroides).[7][78][84][102][121][177][178][179][excessive citations] Occasionally owls are also included in the prey spectrum when an opportunity arises, including barn owls (Tyto javanica), southern boobooks (Ninox boobook) and even powerful owls (Ninox strenua).[7][28][120][180] Wedge-tailed eagles are apex predators
and have no well-documented predators, although presumably they have
some nest predators, likely including ravens and currawongs, especially
when displaced by human disturbance from their nests.[8][9]
Occasionally, these eagles may possibly risk injury or death in
conflicts against other powerful predators and scavengers, such as
dingos, quolls, Tasmanian devils, goannas and snakes, but no such
verified instances seem to be known in literature, and man is considered
to be the wedge-tailed eagle's only true threat.[7][8][9] Occasionally they may be injured and even killed via intra- and interspecies territorial conflicts and mobbing by other birds of prey, especially stooping peregrine falcons
which have successfully knocked wedge-tailed eagles out of the sky,
with a force known to kill both golden and bald eagles in other parts of
the world. Due to the formidable aerial attack of the peregrine, it may
be the only raptor besides the white-bellied sea eagle that
wedge-tailed eagles may avoid nesting near.[8][9][181]
Most of the large falcons, including peregrine, brown and black
falcons, and at times large owls nest in unused or abandoned
wedge-tailed eagle nests.[8][9]
The breeding season is from July to December through much of range, in New Guinea apparently from May on.[4]
They have a distinct tendency that lay earlier in the more northerly
part of the range. For instance, in northeastern Australia laying has
been recorded in January and February and in Tasmania in September.[4][10] In western Australia, breeding depends on food and during drought periods there may be no nesting for up to 4 years.[4]
Adult wedge-tailed eagles are usually solitary or occur in pairs but
immatures are more gregarious. 10–15 young wedge-tailed eagles may rest
or soar together or even hunt together and up to 40 have been recorded
at once at a carcass.[4] Mated adults perform mutual soaring, undulating dives, and tandem flights with rolling and foot-touching.[86][182] The female may appear to ignore or more often turn over and present claws when a male is displaying.[10] As possibly part of courtship feedings on pairs have taken place away from the nest and sharing of a cache of food may occur.[8][183]Allopreening
occurs occasionally between pairs but is seldom observed, although at
times has been considered a “regular” part of the courtship process.[8][86]
Contrary to historical accounts, wedge-tailed eagles seldom engage in
an elaborate courtship display and will instead generally try to
conserve energy, instead devoting their energy for the upcoming trying
breeding season along with territorial exclusions of conspecifics and
obtaining food.[184] Mating tends to occur on a bare branch or dead tree in the nest area, and may continue into the nestling period.[86][185] Contrary to old accounts, the species does not mate in flight.[8]
In the pre-laying phase, mating was recorded to be preceded or
accompanied by loud, slow yelping, but in the nestling period, the pair
alighted together and the male mounted without preliminaries and a
silent copulation lasted for one minute.[7][86]
Territories
are established with aerial displays, which can include high circling
by one or both of pair, sometimes interspersed with flight rolls and
talon presenting.[4]
Most of the time, wedge-tailed eagles typically respect pair boundaries
and can limit territorial behaviour to mild aerial flights, with the
intruders usually giving the ground to incumbents.[8] Violence is usually avoided but sometimes the most heated territorial disputes can escalate to deaths.[8]
Sometimes the displaying eagle may engage in a steep dive on part
closed wings followed by an upwards swoop, later may escalate into
spectacular sky dance with undulations; they may too loop-the-loop.[4][10]
Cartwheeling is typically rare but in one case, three immatures mock
dived at each other, two birds interlocked and cartwheeled several times
before breaking away. No cartwheeling or talon grappling has been
reported between members of a mated pair, but occasionally reported as
used against intruding eagles.[4][182] Aerial displays may go on for a while normally early in the breeding season, between 3 months and 3 weeks prior to egg laying.[10]
Territorial attacks by male wedge-tailed eagles may be against
any encountered intruding eagles, including both male and female
intruders, while female eagles engage in less territorial attacks and
when they do, it is exclusively against other females.[186]
Territorial aggression can extend towards hang gliders and aircraft,
advances noisily, bill open and talons extended until flying just above
and behind or slightly ahead of pilot then swoops repeatedly after
making contact with the hang-glider.[4] A core of some 1 km (0.62 mi) radius around the nest is most fervently defended.[86] Foraging ranges from nest may be up to about 20 km2 (7.7 sq mi).[142] Foraging ranges on breeding home ranges may be around 50 km2 (19 sq mi) for males and 20 km2 (7.7 sq mi) for females in arid central western Australia.[120] Range sizes of pair members vary greatly based on topography, habitat and prey access.[8] Several reported densities of 3–6 pairs per 100 km2 (39 sq mi), others of 7–12 pairs per 100 km2 (39 sq mi).[4]
When rabbits were in plague type numbers, pairs may nest as close 700 m
(2,300 ft) apart and 4 others no more than 2 km (1.2 mi) from those two
pairs.[4] In semi-arid areas of New South Wales near Menindee densities were found to be about a pair per 53 km2 (20 sq mi), 10–12 pairs in good years, 3 in drought years.[187] Not far from that in Mutawintji National Park density was around a pair per 3 to 9 km2 (1.2 to 3.5 sq mi).[54] Much higher densities were noted in this semi-arid zone of western New South Wales, with a pair per 3 to 9 km2 (1.2 to 3.5 sq mi), against around a pair per 40 to 48 km2 (15 to 19 sq mi) in other arid zones.[54] In Western Australia, arid areas had a nearest neighbour distance of 5.32 km (3.31 mi) while those nesting in mesic areas had a distance of 4.88 km (3.03 mi).[188] At Fowlers Gap, there were 9–10 pairs per 390 km2 (150 sq mi).[55][56] Near Canberra, around 37 pairs were reported in an area of 1,000 km2 (390 sq mi), including some unusually as close as 130 m (430 ft) to paved roads
and as close as 260 m (850 ft) from suburban spots. This contrasts
strongly with 36 years prior, when few nests were near human-altered
areas and the amount of pairs in the same area was about 32.[189][161] In the Fleurieu Peninsula in South Australia during the early to mid 2000s, there was a pair per 53 km2 (20 sq mi), active nest sites were 6.6 km (4.1 mi) apart, while the average home range around the nest is 34 km2 (13 sq mi) roughly.[142]
Resurvey efforts a dozen years later in Fleurieu Peninsula found a more
populous population, resulting in a home range estimated at 32.1 km2 (12.4 sq mi) with some active nests as close as 1.5 km (0.93 mi) apart.[190] In the Perth area, it was projected that the mean home range was about 36 km2 (14 sq mi).[28]
Meanwhile, in southern Victoria the nearest neighbour distance of
breeding pairs was 4.7 km (2.9 mi) while mean territory size was
calculated at 17.6 km2 (6.8 sq mi).[191]
Both sexes may participate in building the nest but the female takes
the greater share, often standing in the middle and building outwards.[10]
Often wedge-tailed eagles build alternative nest, up to 2 to 3 per
territory, though when undisturbed uses the same general site
repeatedly.[10] In Tasmania, territories held a mean of 1.4 nests.[8]
The nest is usually either substantial or massive. The nest is a
structure of sticks ranging 70 to 100 cm (2.3 to 3.3 ft) across and 30
to 80 cm (12 to 31 in) deep when first build but with repeated additions
up to 2.5 m (8.2 ft) across and nearly 4 m (13 ft) deep.[4][6] The interior nest cup is commonly around 30 to 40 cm (12 to 16 in) across about 10 cm (3.9 in) deep.[10]
Four studies found the diameter of nests to average from as little as
1.1 m (3.6 ft) and as much as 1.9 m (6.2 ft) and in depth from as little
as 1 m (3.3 ft) to as much as 1.3 m (4.3 ft). Generally speaking in
woodland or forest edge areas, nests tend to be larger, while those in
sparser, more arid areas tend to have characteristically smaller nests,
as they have lesser access to nest building materials.[55][28][56][192]
Good sized nests can weigh well over 400 kg (880 lb).[8] Nests are usually lined with green leaves and twigs, a common practice in accipitrids.[10] Infrequently, they may use an old nest built by another accipitrid, namely whistling kites (Haliastur sphenurus) and white-bellied sea eagles, with the earlier's nests apparently added to in order to enlargen it.[8]
Ideally the nest is located at 12 to 30 m (39 to 98 ft) above the
ground on a lateral branch or main fork of lone or forest tree; in
taller trees, nests can be as much as 75 m (246 ft) high to the opposite
extreme down in lower ground or even on rocks or ground trees are
scarce. In a few studies different areas of New South Wales, the mean
nest height was from 5.2 to 21.7 m (17 to 71 ft) and were often
relatively close to human development.[54][86][55][56][57][excessive citations] Two results in southern Victoria found mean nest heights to be 12.6 and 18.1 m (41 and 59 ft).[191]
In often particularly arid Western Australia, mean nest heights were
reportedly lower, averaging at 6.5 and 15.5 m (21 and 51 ft).[28][120] Detailed study in Western Australia found nest heights were higher in Mediterranean scrubland at 4 to 16 m (13 to 52 ft) against 2 to 6 m (6.6 to 19.7 ft) in the arid zone,
but nest height seemed to not have bearing on occupancy or success,
territoriality kept the population regulated within the habitats.[193] Occasionally they may nest in dwarf trees at as low as 1 to 2 m (3.3 to 6.6 ft).[4][6][193]
Favoured nesting trees include many Eucalyptus and Casuarina species, as well as Corymbia, Callitris and Syncarpia glomulifera while in inland areas more often Acacia, Flindersia as well as Hakea leucoptera and Grevillea striata.[7][86] The amount of Eucalyptus
tree species used by wedge-tailed eagles is extremely diverse and
ultimately the species seem to have no strong overall preferences
regarding tree species, more importantly seeking a given tree of ample
height and considerable broadness.[8][86]
Furthermore, nest trees are often on slightly elevated ground above the
mean ground level, presumably in order to offer a more commanding view
of the surrounding environment.[9][86] Additionally, trees with fewer lower branches may be preferred.[8] Nests are seldom on dead trees, usually this occurs where there is an absence of leafy ones.[7] While Australian nests can be in quite varied surroundings, Tasmanian nests are almost exclusively within well forested areas.[8] Forest type nests tend to have a sparse, open understory and woodlands or nearby glades often are considered perhaps more attractive to the species.[9][86] In desert-type areas, they may nest on a hill or a rise, and in addition sometimes cliff ledges, or among rocks, and even on ground in both islands and desert-like areas, preferably in areas difficult for or inaccessible to humans.[4][6][194] Additionally they've been known to nest on power pylons and telegraph poles.[195][196] Other smaller animals may nest among the sticks at the base of active wedge-tailed eagle nests such as finches, pardalotes and even possums
(which more so than the small birds are presumably vulnerable to the
eagles if caught in the open), perhaps gaining some protection from the
presence of the eagles.[8] This is a not unknown phenomenon in many bird assemblages for small birds to gain incidental protection from strong raptors.[197] Other species, such as Pacific black ducks (Anas superciliosa),
falcons and owls, may also benefit by utilizing unused nests for their
own breeding purposes, although typically only the falcons usually use
them with relative regularity.[8]
Clutch size is usually just one or two but sometimes to 4.[4] About 80% of nests where eagles have managed to lay eggs contain two eggs.[8] Mean clutch size is apparently somewhat higher in the western part of the range.[198] The female lays multi-egg clutches by some 3 days or so apart.[8]
The eggs are buff or white in colour, often appearing heavily blotched
all over with purple-brown, red-brown or lavender, or more sparsely
spotted with reddish brown. The amount of spotting is quite variable on
eggs even within a single clutch, some being heavily marked, others
hardly at all, and at times concentrated on the pointier end of the egg.
When freshly laid, the eggs are glossy but they become more matte and
brittle with age.[8][10]
Eggs may range in height from 66 to 79.3 mm (2.60 to 3.12 in),
averaging 73 mm (2.9 in) in a sample of 54, by 55 to 63.5 mm (2.17 to
2.50 in), averaging 58.8 mm (2.31 in).[8][10][198][199] Each egg normally weighs about 120 to 150 g (4.2 to 5.3 oz), the equivalent to about three chicken eggs or about 3% of the female eagle's body weight, 10% when the clutch number is 3, which is typical for an Aquila eagle but a small percentage relative to smaller raptors.[8][169][198][199] The larger eagles of Tasmania lay larger eggs on average reportedly.[6] Wedge-tailed eagles sometimes lay runt eggs in nests in normal nests, in a condition apparently unique for Australian raptors, and these reportedly never hatch.[8][6]
If a clutch is lost or stolen early in incubation, some pairs have been
documented to replace it, being able to do so about a month later.[8] The incubation stage lasts for 42–48 days.[4] The female of the pair either primarily or entirely incubates on her own and, like many eagles, she is a tight sitter.[8][10][200] However, the male will incubate at times as well, at least up to an hour at a time.[7][121]
In New South Wales, the male was found to incubate for 16–20% of
daylight during which the nest was unguarded for 3–13% of the day.[55][56]
In some cases, male incubation may vary from 1–6% of daylight to as
much as 38% of daylight with shifts in extreme lasting up to 6 hours.[7][86]
Male primarily delivers prey to the nest during incubation (not prior),
up about to the stage where the eaglet(s) can be left unattended.[7][55]
The chicks are covered in white down up at first and are expectedly semi-altricial.
At about 12 days or so, a slightly greyer down develops and this
ultimately becomes the woolly undercoat for the contour feathers. Within
a couple days later, the black quills of the primaries often start to
emerge, and can start to stand and move around the nest. At 28 days, the
eaglets are showing their upper wing coverts increasingly through the
down. At 35 days of age, some darker feathers are appearing on areas
such as the breast, belly, mantle, back and head; mostly these are
evident as a few dark rufous feathers poking through the head down while
at this age they show a short buff-tipped tail. They are partially
feathered up to 37 days and nearly completely feathered by 49 days.[8][10][9][200] At around 37 days, they can attempt to tear food from carcass in the nests without much success.[10]
From 50 days onwards, the eaglet(s) play a good deal, pouncing on
sticks and degree around the nest. Around this age, they are markedly
almost full feathered but for the wing and tail, neither of which has
reached its full length, and they may have a few wisps of down about
their crown or neck. Weight increases are from about 1.17 kg (2.6 lb) at
15 days with a notably increase in robustness to 2.6 kg (5.7 lb) at 29
days, to 3.2 kg (7.1 lb) at 49 days, making more rapid feathering growth
thereafter primarily while body size growth slows considerably.[8][10] Sibling aggression tends to begin at very early stage of life and decrease after first week.[8]
Unlike related eagles, there is some evidence that higher parental
attendance limits instances of aggression, whereas in other eagles this
occurs often in the parent's presence. In all eagles, the parent eagles
do not attempt to intercede when runting or aggression between siblings
occurs.[53][55]Siblicide
occurs occasionally in this species and it is considered a “facultative
cainist” rather than an obligate one, meaning siblicide occurs
occasionally and as conditions dictate as opposed to some eagles where
it occurs almost invariably.[8][53][201] Sufficient environmental conditions can largely reduce sibling aggression.[8]
In New South Wales, three of four successful pairs raised two
fledglings and no sign of rivalry or pecking behaviors observed despite
the size difference of two siblings.[87]
The female broods attentively at first but then decreases after
the second or third week and then ceases brooding almost entirely by 30
days, even at night. For 40 or more days, the female continues to assist
the young with feeding, typically from the male's prey deliveries
though the female may resume hunting after nest attendance drops.[10][202]
Potential predators such as goannas are struck when found to be
approaching the nest, although the eagles usually abandon the nest when a
human approaches. Repeated intrusions and noisy disturbances may have a
net negative effect such as on Tasmanian wedge-tailed eagles, as these
factors often lead to nest failure.[8]
In one case in South Australia, the removal of a dead tree in the
vicinity of a wedge-tailed eagle nest resulted in full abandonment of
the nest by the parents.[182] The female may too continue to bring green leaves to a late stages, doing so more often in a spell of wet weather.[10] During times of plenty, caches can sometimes form around the nest, with much prey left partially or entirely uneaten.[8]
Upon leaving the nest at 11 to 12 weeks of age, the young eagles are
not strong fliers for another 20 days or so, but can competent flying
can be by about 90 days of ages, though full feather development is not
until 120 days.[8][10] Fledgling occurs at 67 to 95 days, typically being less than 90 days and averaging roughly around 79 days.[4][10][55][56][121][excessive citations]
Dependence lasts up to 4 to 6 months after fledging, with the
juvenile eagles which overstay rarely known to be an occasionally fatal
danger to the subsequent chick hatched to their parents.[4][8]
During the later periods of fledgling, interactions are restricted to
brief prey deliveries and the parent eagles stop feeding the young
eagle(s), forcing to go forage elsewhere for foods.[10] A study of post-fledgling dispersal found in one case that a young eagle covered only a 4.22 km2 (1.63 sq mi) range, with a maximum covered in a week of 3.28 km (2.04 mi).[203]
Most recoveries in one banding study were distributed under 300 km
(190 mi) from their original banding site, mostly as fledgling age
juveniles, but some meandered up to 821 km (510 mi) away.[67]
After dispersal, young eagles are floaters up until their 4th or 5th
year, typically avoiding the territories of adults and searching out
feeding opportunities. Up to two-thirds of young wedge-tailed eagles may
die some time between fledgling and when they are 3–5 years but adults
often have quite low mortality rates and can live the better part of a
half century.[8] First breeding is typically at 6 or 7 years old.[193]
Lifespans of wedge-tailed eagles in the wild are poorly known, with the
maximum recorded in one banding study being merely 9 years, quite a
paltry age compared to other large eagles, and it is quite conceivable
that eagles who survive to maturity not infrequently live around twice
that long or more. In captivity, the species has been known to live to
around 40 years of age.[8][9]
Only one young is typically produced from a clutch of two but
occasionally two fledglings may occur. The breeding success rates of
the species are variable.[6]
In overall studies, at least 52 to 90% of breeding pairs managed to
produce a fledgling, with further projected numbers from this of 0.2–0.5
fledgling per pair, 0.7–1.2 fledgling per clutch and 1.1–1.3 fledgling
per brood.[8] In southwestern Australia, from 0.7 to 1.2 young are fledged per clutch laid, 0.19–0.46 young per pair per year.[6] In south-central Queensland, fledgling productivity was 1.1 per young to pairs that laid eggs.[95]
Northern New South Wales eagles were able to produce 0.8 young per pair
from 2005 to 2006 while 0.89 and 0.64 fledged young per pair per year
was the fledgling rate in central and western New South Wales,
respectively.[86][55][204] In a further study in New South Wales at Burrendong Dam,
from 1993 to 2003, 15 pairs produced an average of one fledgling per
territory but 1998 due to drought conditions, the rate was only 0.4
chick per territory. Within Kinchega National Park, however, the rate of 0.99 young per pair was fairly consistent regardless of climatic conditions.[8] In the Australian Capital Territory, pairs were said to produce 1.1 fledglings per pair.[189]
In southeastern Australia, from 0.9 to 1.5 young per clutch are laid,
with 0.6–1.0 young per pair per year. In Tasmania, from 0.64 to 0.8
young are fledged per clutch laid, 1.07 per successful nest.[7][6] 0.91 young were produced per pair in southern South Australia or 1.1 fledglings per successful nesting attempt.[142]
Subsequent research in South Australia found 38 successfully
fledged young with 10 pairs or 26% producing two fledglings and that
production was 1.1 per occupied territory and 1.3 per successful pair.[205] 0.73 fledglings were produced pair per year in south-west Western Australia.[28] Western Australian eagles produced 0.92 fledged young per clutch laid and 1.1 young per successful nest.[206] During periods of drought in Western Australia, some wedge-tailed eagles may forgo breeding for up to four years.[6][207]
Higher annual rainfall in Western Australia, higher in mesic than arid
areas, made a big difference in pair productivity, with 12% of arid zone
pairs producing young, or 0.13 fledglings per pair, a very low
productivity, while the mesic zone 69% of pairs produced fledglings, or
0.77 fledglings per pair.[188]
Generally, wedge-tailed eagles can nest in a variety of habitats and
climatic conditions but tend to be slightly less productive in more arid
environments.[207]
Significant broad-scale control is thought to be unlikely to be harming
numbers of young being produced with those with a macropod-based diet
perhaps having a richer diet.[7][86] Like most eagles, wedge-tailed eagles fit the mould rather well of a K-selected breeder, i.e. being large, producing fewer young and tending to live relatively long.[207]
In the 1990s, it was estimated broadly that the global population was somewhere between 10,001 and 1,000,000 individuals.[4][15] As of 2009, Birdlife International listed the total population as only 100,000 mature individuals, possibly conservative and from admittedly poor supporting data.[1][15][208] As of that analysis, Birdlife considers the overall population of wedge-tailed eagles to be “possibly increasing”.[1] Generally, the wedge-tailed eagle appears to be quite stable in population.[15][209]
Although wedge-tailed eagles are often scarcer than those large
distribution suggests, their total distribution covers more than 10.5
million square kilometres and the population is quite likely within
hundreds of thousands.[1][4] Thinning of forest cover,
mostly inadvertent provisioning of carrion food sources and,
particularly, rabbit introductions may have aided the species, and it
may actually be commoner now than before European colonization.[4][6]
Though protected, sometimes wedge-tailed eagles are shot or trapped or
killed by poison carcasses set out by farmers many of whom consider it a
serious sheep killer.[4] Historically, the wedge-tailed eagle was subject to persecution levels to rival any other eagle in the world.[10][210][12]
The heavy persecution began in the closing decades of the 19th century,
due largely to the establishment of large-scale sheep farming in
Australia. One Queensland station claimed to have poisoned 1060 eagles
over 8 months in 1903.[8]
Laws passed from 1909 to 1925 made it mandatory for landowners
and farmers to kill eagles as vermin with enforcement determined by a
given region's minister or vermin board, resulting in even more sweeping
efforts to destroy the species.[12][211]Steel-jawed rabbit traps were set around carcasses and Heligoland traps could sometimes trap several eagles at once, beyond sustained shooting and poisoning efforts.[8]
Between the years 1958 and 1967, 120,000 bounties were paid in merely
the states of Queensland and Western Australia on wedge-tailed eagles
killed, meaning an average 13,000 were killed each year.[8][12][211]
Even by 1967 to 1976, likely intentional human killings accounted for
54% of wedge-tailed eagle mortalities in Western Australia, with an
estimated 30,000 killed in the year of 1969 throughout Australia.[8]
Strong legal protections started in Western Australia in the 1950s
increasingly so to the 1970s or later elsewhere, now it is protected and
subject to limited persecution throughout.[12] Despite reduced persecution, as of the 1980s, 54% of recovered eagles by the 1980s were killed by human persecution.[212]
Despite such stunningly high rates of persecution, the wedge-tailed
eagle was remarkably resilient to the haphazard persecution inflicted by
humans in a way many other Australian wildlife, especially the
regionally endemic mammals, and even other eagles elsewhere often are
not.[25][213][214]
Often the species is less intentionally harmed via human disturbance via land development
particularly intensifying agricultural and modern settlements, which
can in turn lead to clearing of mature trees, disturbances at the nest
and decline of native prey species, all of which have a net negative
effect on the wedge-tailed eagles.[7][6][215] Eggshell thickness was not significantly decreased by the use of DDT
likely due likely to the largely mammal-based diet of the species,
whereas raptors which consume birds or fish are disproportionately
effected by DDT.[6][216] On occasion, the species is still subject to illegal shootings and poisonings, however persecution of the species is significantly less prevalent in recent decades.[8][6][67] Occasionally but not commonly, they are killed by sodium fluoroacetate
poisons long used to “control” Australian wildlife, but now generally
directed at invasive species such as rabbits, feral pigs and foxes.[8][9]
A list of the main persistent threats in the 21st century to
wedge-tailed eagles consists of: destruction of habitat, including logging, developments including urbanization, wind farm collisions and the disturbance and destruction associated with their construction, increasing density rural human populations, illegal persecution in sheep farm areas, drowning in open tanks in drypastoral zones, roadkills (especially while foraging for roadkill carrion), collisions with fences, powerlines and airplanes, regular electrocutions, poisoning from rabbit baits and other baits and exposure to lead and other bullet fragments which may be responsible for some eagle debilitations and deaths.[7][217][218][219][220][excessive citations] Within the Fleurieu Peninsula, some 1.74 eagles on average are claimed by wind farm turbine collisions.[142]
Conservation needs may differ in different habitats, i.e. in more
coastal temperate areas, the eagle is reported to have difficulty
nesting when hillsides have been cleared of trees, meanwhile inland,
they have lesser need of trees in elevated locations because they are
more often assisted by thermals. However, they cannot generally persist where leafy trees are clearcut.[8][221] Surprising resilience even to drought was found recently in the wedge-tailed eagles in the Australian Capital Territory
where pair occupancy remained consistent through drought for
wedge-tailed eagles but not for little eagles, but this is may have more
to do with the wedge-tails more successful uncoupling from a dependence
on declining rabbits as prey than the little eagle.[222]
Of 84 eagle deaths or debilitating injuries, 52% were attributable to
collisions or electrocutions, 15.5% due to persecution, 11% due to
natural causes and 15% were due to unknown causes.[8]
The Tasmanian race of wedge-tailed eagle, A. a. fleayi, is
quite restricted in range and habitat, with estimated numbers having
gone from 140 pairs in the 1980s down to only 60–80 by the mid-1990s.[4][6][223]
With the island's population numbering quite low and likely continuing
its declining, as evidenced by slow replacement of lost pair members,
the subspecies is listed as state-endangered.[8][224][225]
Furthermore, surveys contrasting 1977–1981 with 1998–2001 data found a
decline of around 28% in the island's reported number of eagles.[7]
Generally Tasmanian wedge-tailed eagles are even less tolerant of human
alterations and disturbances near the nest site than mainland
wedge-tailed eagles and have more specific habitat requirements.[4][6] Historically, the same hunting organization in Tasmania that played a large role in the extinction of the thylacine (Thylacinus cynocephalus)
also intentionally tried to hunt the Tasmanian wedge-tailed eagle into
extinction, publicly having erroneously claimed that eagles were
non-native in Tasmania; however, hunting is unlikely to further continue
on a large scale in the state.[8]
Where habitat clearance and degradation is extensive in Tasmania, the
native prey populations are insufficient to support eagles. Furthermore,
the clearing or logging of trees is especially critical in Tasmania,
where the eagle is by and large a forest-dependent breeder.[8][192][226]
Studies indicate that Tasmanian eagles mostly nest in emergent
trees in old-growth native forest exposed to early morning sun and
sheltered from prevailing strong winds and cold spring winds, given the
more temperate climate there relative to most points in mainland
Australia.[8][192][227]
The subspecies requires forest areas greater than 10 ha (25 acres) in
which to breed and is very prone to desert its nest when disturbed.[8][192] A predicted change was calculated to the carrying capacity of the Tasmanian forest given current operations is modeled, likely driving the population down.[11]Wind farms
in Tasmania are also an occasional threat; although not thought to be a
significant source of mortality, wedge-tailed eagles, especially young
ones, are less success at avoiding invariably fatal collisions with them
than Tasmanian white-bellied sea eagles.[228][229] Furthermore, of 109 eagle carcasses recovered in Tasmania, all of them had trace levels of lead in their livers or femurs with at least part of the exposure likely from lead ammunition.[230]
In addition, like all eagles, Tasmanian wedge-tailed eagles are
vulnerable to electrocutions, and collisions with vehicles, overhead
wires, and fences and poisonings, largely via illegal killings by
poachers of Tasmanian devils and forest ravens (Corvus tasmanicus).[15][192][231] Efforts are underway to ameliorate the harm being done to Tasmanian wedge-tailed eagles, especially via forestry operations.[231]
In protected areas,
protocols are in place to protect Tasmanian eagle nests and protect
them by creating an obligatory nest reserve of at least 10 ha and forestry
operation have been restricted during the breeding season to outside a
buffer zone of 500 m (1,600 ft), extending further to 1 km (0.62 mi) if
the proposed work is in the line-of-sight of the nesting eagles.[8][231][232]
About 20% of known pairs are outside protected areas and on private
land, so are largely outside the strict legal protection the subspecies
has on governmental forest land.[15][231]
Furthermore, researchers are instituting rules to minimize disturbance,
limiting breeding surveys to distant observations of whitewash and
flattened treetops as proof of nesting and all detailed observations to
be obtained after the cessation of breeding activities.[192]
Early in 1967, the Australian Army 2nd Cavalry Regiment
received its new badge, a wedge-tailed eagle swooping, carrying a
lance-bearing the motto "Courage" in its talons. The regiment's mascot
is a wedge-tailed eagle named "Courage". Since its formation, there have
been two, Courage I and Courage II. In 1997, while on flight training
with his handlers, Corporal Courage II refused to cooperate and flew
away, not being found for two days following an extensive search. He was
charged with being AWOL and reduced to the rank of trooper. He was
promoted back to corporal in 1998.[235][236]
The West Coast Eagles, an AFL football club from Western Australia,
uses a stylised wedge-tailed eagle as their club emblem. In recent
years, they have had a real-life wedge-tailed eagle named "Auzzie"
perform tricks before matches.[23
Allée Des Baobabs near Morondava, Madagascar
At 
An adult wedge-tailed eagle at 
Taking off from its perch, the long legs of this adult female are clearly visible.
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